The Combining Greek Ypogegrammeni at U+0345 is a nonspacing mark that adds the small “iota under” sign used in polytonic Greek text. It belongs to the Combining Diacritical Marks block, and it attaches to a preceding Greek vowel rather than standing as an ordinary visible letter. Its main use is in Ancient Greek and scholarly editions of Greek, where it represents the historical iota subscript written under long vowels such as alpha, eta, and omega. In words such as τῇ or ᾳ, it marks an old iota that affected historical spelling and grammar but is not pronounced as a separate vowel in standard classical reading traditions. Editors, classicists, theologians, lexicographers, and digital libraries use it when preserving full polytonic orthography in quotations, critical texts, dictionaries, and inscriptions transcribed into modern type. It commonly appears alongside Greek accents and breathings, so the same base vowel may carry several combining marks at once. The practical trap is normalization: many common Greek characters with iota subscript also exist as precomposed letters, so visually identical text may be stored in different ways. That can affect searching, sorting, and text comparison unless the software treats canonical equivalents consistently.